Interesting thing here is that I think any of these scientists could have drawn parallels to human instinct to murder, and the topic would have been much less controversial.
Other animals practice cannibalism, even of their own young, or violence towards family members, etc. I'm not arguing whether those behaviors are more or less repellent than rape, but I would suggest that making clearer the attempt to draw parallels to other animals might help to defuse some of the (understandable) uproar.
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